The Harlequin Ladybird Survey
Harlequin ladybird elytra
HOME What is a ladybird?Recognising the Harlequin ladybirdFact FileResearchRecording sightingsFor young peopleAcknowledgementsContactsUseful links
Harlequin Ladybird Project Publications

Brown, P.M.J. & Roy, H.E. (2007) The spread of the Harlequin Ladybird Harmonia axyridis in late 2006. Atropos 31: 32-33.

Brown, P.M.J. (2007) In love with ladybirds. BBC Wildlife 25(7): 58-63.

Brown, P.M.J., Adriaens, T., Bathon, H., Cuppen, J., Goldarazena, A., Hagg, T., Kenis, M., Klausnitzer, B.E.M., Kovar, I., Loomans, A.J., Majerus, M.E.N., Nedved, O., Pedersen, J. Rabitsch, W., Roy, H.E., Ternois, V., Zakharov, I. & Roy, D.B. (2008) Harmonia axyridis in Europe: spread and distribution of a non-native coccinellid. BioControl 53: 5-22.

Brown, P.M.J., Roy, H.E. & Majerus, M.E.N. (2007) The spread of the harlequin ladybird in Cambridgeshire and Huntingdonshire. Nature in Cambridgeshire 49: 55-57.

Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (2006) Ladybird recording in Cambridgeshire and Huntingdonshire. Nature in Cambridgeshire 48: 12-16.

Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (2007) Update on the spread of an invasive ladybird. Biocontrol News and Information 28(1): 16-17.

Brown, P.M.J., Roy, H.E., Rothery, P., Roy, D.B., Ware, R.L. & Majerus, M.E.N. (2008) Harmonia axyridis in Great Britain: analysis of the spread and distribution of a non-native coccinellid. BioControl 53: 55-68.

Brown, P.M.J., Roy, H.E., Ware, R.L. and Majerus, M.E.N. (2006) Distribution and identification of the harlequin ladybird Harmonia axyridis. Atropos 28: 63-69.

CAB International (2006) Harmonia axyridis [original text by H.E. Roy, P.M.J. Brown and M.E.N. Majerus]. Crop Protection Compendium, 2006 edition. Wallingford, UK: CAB International.

Hall, R., Ware, R.L., Michie, L.J., Brown, P.M.J. and Majerus, M.E.N. (submitted) First occurrence of Rhyzobius chrysomeloides (Herbst, 1792) (Coleoptera: Coccinellidae) in Cambridgeshire. Entomologistís Record and Journal of Variation.

Holroyd, O., Brown, P.M.J., Roy, H.E. and Majerus, M.E.N. (in press) The harlequin ladybird, Harmonia axyridis, reaches Scotland. Entomologistís Record and Journal of Variation.

Karyistinou, A., Thomas, A.P.M. & Roy, H.E. (2004) Presence of haemocyte-like cells in coccinellid reflex blood. Physiological Entomology 29: 94-96.

Kenis, M., Brown, P.M.J., Ware, R.L. and Roy, D.B. (in press) Invasion of the harlequin ladybird, Harmonia axyridis, in Europe: when beauty becomes the beast. ALARM Atlas of Biodiversity Risks.

Majerus, M.E.N. and Roy, H.E. (2005) Arrival of the Harlequin Ladybird in Britain. Biological Sciences Review, Sept. 2005: 29-32.

Majerus, M.E.N. and Roy, H.E. (2005) Scientific opportunities presented by the arrival of the Harlequin ladybird, Harmonia axyridis, in Britain. Antenna 29(3): 196-208.

Majerus, M.E.N., Brown, P.M.J., Roy, H.E. and Ware, R.L. (2009) Ladybirds (Wildlife reports). British Wildlife 20: 210-212.

Majerus, M.E.N., Mabbott, P., Rowland, F. and Roy, H.E. (2006) The harlequin ladybird, Harmonia axyridis, arrives in Britain. Entomologistís Monthly Magazine 142: 87-92.

Majerus, M.E.N., Roy, H.E., Brown, P.M.J., Ware, R.L. and Shields, C. (2006) A Guide to the Ladybirds of the British Isles (fold-out chart). Field Studies Council, Shrewsbury.

Majerus, M.E.N., Strawson, V. & Roy, H.E. (2006) The potential impacts of the arrival of the Harlequin ladybird, Harmonia axyridis (Pallas) (Coleoptera: Coccinellidae), in Britain. Ecological Entomology, 31: 207-215.

Michie, L.J., Disney, R.H.L., Hall, R.J., Ware, R.L. & Majerus, M.E.N., (in press). First occurrence of Phoridae (Diptera) and live Harmonia axyridis (Pallas) (Col.: Coccinellidae) eclosing from the same pupae. The Entomologistís Record

Pascoe, S., Brown, P.M.J. and Majerus, M.E.N. (2007) Hybrid mating between Harmonia axyridis and Adalia bipunctata. Bulletin of Amateur Entomologists' Society 66:156-159.

Pell, J. K., Baverstock, J., Roy, H. E., Ware, R. L. and Majerus, M. E. N. (2008). Intraguild predation involving Harmonia axyridis: a review of current knowledge and future perspectives. BioControl 53: 147-168.

Rhule, E. L., Majerus, M. E. N. & Ware, R. W. (in prep) Potential use of the sexually-transmitted mite Coccipolipus hippodamiae (Acaridae: Podopolipidae) in controlling populations of the invasive ladybird Harmonia axyridis (Coleoptera: Coccinellidae)

Ribbands, B., Brown, P.M.J., Roy, H.E. & Majerus, M.E.N. (in press) The most northerly record of the harlequin ladybird in the British isles. Entomologistís Monthly Magazine.

Roy, H. E., Baverstock, J., Ware, R. L., Clark, S. J., Majerus, M. E. N., Baverstock, K. E. & Pell, J. K. (2008). Intraguild predation of the aphid pathogenic fungus Pandora neoaphidis by the invasive coccinellid Harmonia axyridis. Ecological Entomology 33: 175-182.

Roy, H.E. and Majerus, M.E.N. (2005) Unlovely Ladybirds? Biological Sciences Review, 17(4): 42.

Roy, H.E.& Cottrell, T. (2008) Forgotten natural enemies: interactions between coccinellids and insect-parasitic fungi. European Journal of Entomology, 105: 391-398.

Roy, H.E., Baverstock, J., Ware, R.L., Clark, S.J., Majerus, M.E.N., Baverstock, K.E. & Pell, J.K. (2008). Intraguild predation of the aphid pathogenic fungus Pandora neoaphidis by the invasive coccinellid Harmonia axyridis. Ecological Entomology, 33: 1-8.

Roy, H.E., Brown, P.M.J. and Majerus, M.E.N. (2006) Harmonia axyridis: A successful biocontrol agent or an invasive threat? In: An ecological and societal approach to biological control (eds.: J. Eilenberg & H. Hokkanen). © Kluwer Academic Publishers, Netherlands.

Roy, H.E., Brown, P.M.J., James, T.J., Munford, J. and Majerus, M.E.N. (2005) Monitoring an Alien: Harmonia axyridis. Journal of Practical Ecology and Conservation Special Series, No.4: 77-82.

Roy, H.E., Brown, P.M.J., Rothery, P., Ware, R.L. & Majerus, M.E.N. (2008) Interactions between the fungal pathogen Beauveria bassiana and three species of ladybird: Harmonia axyridis, Coccinella septempunctata and Adalia bipunctata. BioControl 53: 265-276.

Roy, H.E., Rowland, F., Brown, P.M.J., and Majerus, M.E.N. (2005) Ecology of the harlequin ladybird: a new invasive species. British Wildlife 16(6): 403-407.

Ware, R.L. & Majerus, M.E.N. (2008) Intraguild predation of immature stages of British and Japanese coccinellids by the invasive ladybird Harmonia axyridis. BioControl, 53: 169-188.

Ware, R.L., Evans, N., O'Farrell, K., Malpas, L., Michie, L.J. & Majerus, M.E.N. (2008) Intraguild predation of British and Japanese coccinellid eggs by the invasive ladybird Harmonia axyridis. Neobiota, 7: 263-275.

Ware, R.L., Majerus, M.E.N., Roy, H.E. and Symington, F. (2005). The harlequin ladybird arrives in Britain: a threat to our native species? Bulletin of the Amateur Entomologists' Society 64: 175-186

Ware, R.L., Ramon-Portugal, F., Magro, A., Ducamp, C., Hemptinne, J-L. & Majerus, M.E.N. (2008) Chemical protection of Calvia quatuordecimguttata eggs against intraguild predation by the invasive ladybird Harmonia axyridis. BioControl, 53: 189-200.

Ware, R.L., Yguel, B. & Majerus, M.E.N. (2008) Effects of larval diet on female reproductive output of the European coccinellid Adalia bipunctata and the invasive species Harmonia axyridis. European Journal of Entomology, 105: 437-443.

Ware, R.L., Yguel, B. & Majerus, M.E.N. (2009) Effects of intraguild predation and competition on larval development of the European coccinellid Adalia 2-punctata and the invasive species Harmonia axyridis. Ecological Entomology 34: 12-19

Top